Of course, wed like these snails to reproduce and they have proved to be able to do so in captivity. As front-gilled snails they are dioecious, the male fecundating the female by passing a spermatophore. This is a lump of spermatozoa held together by cement liquids produced by glandular appendages of the sexual organs.
The offspring grow in eggs in the mothers brood pouch; each embryo lying in its own section and feeding on the nourishing substance there.
When the embryos development is sufficiently advanced it is transported to the outside from under the mother snails shell lip in a white-ish egg. Within a few seconds the white substance covering the juvenile dissolves and a fully developed youngster emerges.
The egg is transported to the outside in a groove in the mother snails right side. Depending on the species, the juvenile snail is 0.28-1.75cm/0.1-0.7 long.
When the snail has recently been introduced to an aquarium, so-called shock births happen frequently, possibly caused by differences in the water parameters so snailkeepers are able to watch these births live shortly after putting the snail into their tanks.
The juveniles born inside their eggs are often a little smaller than usual. However, they can survive but under normal circumstances they would just have remained inside the mother snails brood pouch a little longer.
Generally, larger species release larger and less juveniles than the smaller Tylomelania.
These snails do not produce a massive population, as they usually release just one young at a time at intervals, and these juveniles need a long time to grow from only a few millimetres to a respectable size.